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Urtica dioica

L.

Stinging nettle, Greater Nettle

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Urtica dioica, often known as common nettle, burn nettle, stinging nettle, nettle leaf, or just a nettle or stinger, is a herbaceous perennial flowering plant in the family Urticaceae. Originally native to Europe, much of temperate Asia and western North Africa, it is now found worldwide. The species is divided into six subspecies, five of which have many hollow stinging hairs called trichomes on the leaves and stems, which act like hypodermic needles, injecting histamine and other chemicals that produce a stinging sensation upon contact ("contact urticaria", a form of contact dermatitis). The plant has a long history of use as a source for traditional medicine, food, tea, and textile raw material in ancient (such as Saxon) and modern societies.

Description

A herb that continues to grow year after year. It can be 1-2 m high. The leaves have stalks. They are opposite. The leaves are 4-15 cm long by 1-7 cm wide. They have a leaf like structure at the base of the leaf stalk. The leaves are pointed. The flowers are white or yellow. They have male and female flowers on separate plants. The plant has stinging hairs on the leaves and stems.

Edible Uses

Young leaves are cooked as a potherb or added to soups, and can also be dried for winter use. Nettles are highly nutritious, easily digested, and rich in minerals (especially iron) and vitamins (especially A and C). Harvest only young leaves — old leaves can be laxative — and wear stout gloves. Cooking or thorough drying neutralises the sting. Young shoots harvested in spring when 15–20 cm long, complete with the underground stem, are particularly good eating. The leaves are harvested commercially for chlorophyll extraction, used as green colouring agent E140 in foods and medicines. A tea made from the dried leaves is warming and, though bland on its own, can be added as a tonic to China tea. The juice of the leaves or a decoction of the herb can be used as a rennet substitute to curdle plant milks. Nettle beer is brewed from the young shoots.

Traditional Uses

The tender shoots and leaves are cooked and eaten as a vegetable. They can be used in soups and pasta dishes. The leaves are used for sarma in Turkey. They are rolled around a filling of rice or minced meat. The juice of the leaves is used as a rennet in preparing cheeses and junkets. The dried leaves are used as tea. The seeds yield an edible fatty oil. CAUTION: The older leaves can contain oxalate crystals. Raw leaves have stinging properties.

Medicinal Uses

Nettles have a long history as a home herbal remedy and nutritious food. Leaf tea has traditionally been used as a cleansing tonic and blood purifier, and the plant is used in treating hay fever, arthritis and anaemia. The whole plant is antiasthmatic, antidandruff, astringent, depurative, diuretic, galactogogue, haemostatic, hypoglycaemic and a stimulating tonic. An infusion is valuable for stemming internal bleeding and treating anaemia, excessive menstruation, haemorrhoids, arthritis, rheumatism and skin complaints including eczema. Externally it is applied to arthritic pain, gout, sciatica, neuralgia, haemorrhoids and hair problems. Fresh leaves rubbed or beaten onto the skin treat rheumatism through urtification — the intense irritation acts as a counter-irritant, drawing blood to the area to help clear rheumatic toxins, while the formic acid is thought to benefit affected joints directly. For medicinal use, the plant is best harvested in May or June as it comes into flower and dried for later use. Nettle leaf juice can antidote nettle stings, and an infusion of fresh leaves makes a soothing lotion for burns. The root has shown beneficial effects on enlarged prostate glands. A homeopathic remedy made from the leaves treats rheumatic gout, nettle rash and chickenpox, and is applied externally to bruises. The German Commission E approves Urtica dioica leaf for rheumatic ailments (internal use) and irrigation therapy, and the root for inflammatory disease of the lower urinary tract, prevention of kidney gravel formation, and urination difficulty from benign prostatic hyperplasia. The species merits further study for potential uses against kidney and urinary system ailments.

Known Hazards

The leaves of the plants have stinging hairs, causing irritation to the skin. This action is neutralized by heat or by thorough drying, so the cooked leaves are perfectly safe and nutritious. However, only young leaves should be used because older leaves develop gritty particles called cystoliths which act as an irritant to the kidneys. Possible interference with allopathic drugs for diabetes mellitus, hypertension. Central nervous system depression drugs (e.g. morphine, alcohol) may also interact with nettle. Avoid during pregnancy.

Distribution

It is a temperate plant. In Nepal they grow from 500-4500 m altitude. They grow in moist areas. Garhwal Himalayas. It suits hardiness zones 3-9.

Where It Grows

Africa, Alaska, Albania, Andorra, Armenia, Asia, Australia, Austria, Balkans, Belarus, Belgium, Bhutan, Bosnia, Brazil, Britain, Bulgaria, Canada, Caucasus, Central Africa, Central Asia, Chile, China, Congo DR, Croatia, Czech Republic, Estonia, Europe*, Finland, France, Georgia, Germany, Himalayas, Hungary, India, Iran, Italy, Kazakhstan, Kyrgyzstan, Latvia, Lebanon, Lesotho, Libya, Lithuania, Luxembourg, Macedonia, Mediterranean, Middle East, Moldova, Morocco, Netherlands, New Zealand, Nepal, North Africa, North America, Northeastern India, Norway, Pakistan, Poland, Portugal, Romania, Russia, Scandinavia, Serbia, Siberia, Sicily, Sikkim, Slovakia, Slovenia, South Africa, Southern Africa, South America, Spain, Sweden, Switzerland, Tajikistan, Tasmania, Tibet, Turkey, Türkiye, Ukraine, USA, Uzbekistan,

Cultivation

Prefers a soil rich in phosphates and nitrogen. Plants must be grown in a deep rich soil if good quality fibre is required. Nettles are one of the most undervalued of economic plants. They have a wide range of uses, for food, medicines, fibres etc and are also a very important plant for wildlife. There are at least 30 species of insects that feed on it and the caterpillars of several lepidoptera species are dependant upon it for food. Especially when growing in rich soils, the plant can spread vigorously and is very difficult to eradicate. It is said that cutting the plant down three times a year for three years will kill it. It is a good companion plant to grow in the orchard and amongst soft fruit. So long as it is not allowed to totally over-run the plants, it seems to improve the health of soft fruit that grows nearby and also to protect the fruit from birds, but it makes harvesting very difficult. Dioecious. Male and female plants must be grown if seed is required. The leaves and young shoots are typically harvested in spring, before flowering, from early spring to late spring. Stinging nettle usually flowers from late spring to summer, around June to August (Northern Hemisphere). Stinging nettle grows quickly and can reach its full height within a single growing season, making it a vigorous and robust plant.

Propagation

Sow seed in spring in a cold frame, barely covering it. Prick seedlings into individual pots when large enough to handle and plant out in summer. Division succeeds at almost any time in the growing season — plant divisions straight into their permanent positions.

Other Uses

Nettle can serve as ground cover that prevents soil erosion and improves soil quality through nutrient accumulation, while also attracting beneficial insects. A strong flax-like fibre from the stems — produced in less abundance than flax and more difficult to extract — is used for string, cloth and good quality paper, harvested as the plant dies down in early autumn and retted before extraction. Remaining plant matter provides biomass for manufacturing sugar, starch, protein and ethyl alcohol. Seed oil is used as an illuminant. The plant is an essential ingredient of 'QR' herbal compost activator, a dried powdered herbal blend that speeds bacterial activity in compost heaps. Leaves are a valuable compost addition and can be soaked for 7–21 days in water to make a nutritious liquid plant feed that also acts as an insect repellent and foliar feed. The growing plant raises the essential oil content of neighbouring plants, boosting their resistance to insect pests. Flies are repelled by nettles, and freshly cut stems placed in food cupboards exploit this effect. Plant juice or a decoction boiled in strong salt solution curdles milk as a rennet substitute; rubbed into leaky wooden tub seams it coagulates and seals them. Infused leaves make a hair wash used as a tonic and antidandruff treatment. A permanent green dye comes from a decoction of leaves and stems; a yellow dye from the root boiled with alum. Nettle flowers are attractive to pollinators and provide nectar and pollen. The leaves provide food and habitat for caterpillars and other insects, and the dense growth offers shelter for various invertebrates. Nettle is a dynamic accumulator, gathering minerals from the soil and storing them in bioavailable form useful as fertilizer or mulch improver.

Other Information

It is a commercially cultivated vegetable. It is a popular food in hilly regions. It is sold in local markets and commercial shops.

Notes

Nettles especially the roots have been found to be useful for reducing prostrate cancer problems in men. Rich in polysaccharides, vitamins (β-carotene), iron, potassium, manganese, calcium, silicon, phosphate and vitamin C. Its leaves contain a large amount of essential fatty acids, such as α-linolenic acid and linoleic acid, essential amino acids and carbohydrates, carotenoids that include Omega-6 fatty acids, lutein, β-carotene. 75% of the total chlorophyll content is α-chlorophyll and 25% β-chlorophyll.

Nutrition

PartMoisturekJkcalProteinVit AVit CIronZinc
Leaves167407.4
Leaves89157381.8
Leaves dry79.5436.9

Synonyms

Urtica galeopsifolia Wierzb. ex Opiz

Also Known As

Achoka, Ain, Aluma, Ardica, Bichhu booti, Bichhua, Bichua, Bichubuti, Bicirgan, Bobatsi, Bolodzi makholo, Brandnetel, Brennesle, Brennnessel, Chichru, Ch'inch'ari, Chitchiti odghin, Dalgan, Deli isirgan, Derdinik, Dilgeles, Dzaluma, Eghinj, Gasgask, Gazada ut, Gazanga, Gerzinik, Gesges, Gewone brandnetel, Gezik, Gezink, Gezgezok, Gezok, Gharya shisnu, Gicirgan, Gizirkan, Hitha, Horriga lharcha, Horriga lharra, Hourrigua-iharcha, Irhawurhawu, Isiran, Isirgan, Isirganotu, Jamila, Jam jama, Jinch'ari, Jula, Juon, Kabarcik, Kalele, Kandali, Kari, Keyori, Kirays, Kopriva, Korvenoges, Krapiva, Kungsh, Kushak, Lardica sarvaggia, Mahlolo-abolodzi, Noges, Ortia, Ortiga, Ourtya, Patle sishnu, Pokriva, Pokrzywa, Polo, Polu, Raudnoges, Saag, Saddar, Sa-dug, Sadukpa, Sagoe, Sajilim, Sebabetsane, Sezonay, Sezonkay, Seezunkay, Sha zhi, Shouzhe, Siiloi, Sirgan, Sishnu, Sisni, Sisno, Sisnu, Sungri-surong, Tall nettle, Ububati, Urtiga, Urtio, Urzica, Velika kopriva, Xia di, Yagij, Za god, Zahoye, Zara, Zastisod, Ziiloi, Zocha, Zubuda, Zwa

References (192)

  • Agarwal, R. & Chandra, V., 2019, Diversity of wild edible plants in the Mandal-Chopta forest, Uttarakhand. Journal of Medicinal Plants Studies 2019; 7(1): 89-92
  • Aipeisova S.A., et al, 2019, Wild useful herbs of Aktobe Region (Western Kazakhstan). Ukrainian Journal of Ecology, 9(3), 329-331
  • Ali, H., et al, 2011, Ethnobotanical profile of some plant resources in Malam Jabba valley of Swat, Pakistan. Journal of Medicinal Plants Research Vol. 5(18), pp 4676-4687
  • Ambasta, S.P. (Ed.), 2000, The Useful Plants of India. CSIR India. p 663
  • Amin, M., et al, 2023, Edible wild plant species used by different linguistic groups of Kohistan Upper Khyber Pakhtunkhwa (KP), Pakistan. Journal of Ethnobiology and Ethnomedicine 19:6 p 14
Show all 192 references
  • Ari, S., et al, 2015, Ethnobotanical survey of plants used in Afyonkarahisar-Turkey. Journal of Ethnobiology and Ethnomedicine 11:84
  • Aryal, K. P. et al, 2009, Uncultivated Plants and Livehood Support - A case study from the Cheopang people of Nepal. Ethnobotany Research and Applications. 7:409-422
  • Aryal, K. P., et al, 2018, Diversity and use of wild and non-cultivated edible plants in the Western Himalaya. Journal of Ethnobiology and Ethnomedicine (2018) 14:10
  • Awan, M. R., et al, 2011, Studies on traditional knowledge of economically important plants of Kaghan Valley, Mansehra District, Pakistan. Journal of Medicinal Plants Research Vol. 5(16), pp. 3958-3967
  • Beckstrom-Sternberg, Stephen M., and James A. Duke. "The Foodplant Database." http://probe.nalusda.gov:8300/cgi-bin/browse/foodplantdb.(ACEDB version 4.0 - data version July 1994)
  • Bhatia, H., et al, 2018, Traditionally used wild edible plants of district Udhampur, J&K, India. Journal of Ethnobiology and Ethnomedicine (2018) 14:73
  • Bhattarai, S and Chaudary, R. P., 2009, Wild Edible Plants Used by the People of Manang District, Central Nepal. Ecology of Food and Nutrition, 48:1-20
  • Biscotti, N. & Pieroni, A., 2015, The hidden Mediterranean diet: wild vegetables traditionally gathered and consumed in the Gargano area, Apulia, SE Italy. Acta Societatis Botanicorum Poloniae 84 (3): 327-338
  • Biscotti, N. et al, 2018, The traditional food use of wild vegetables in Apulia (Italy) in the light of Italian ethnobotanical literature. Italian Botanist 5:1-24
  • Blamey, M and Grey-Wilson, C., 2005, Wild flowers of the Mediterranean. A & C Black London. p 35
  • Blanco-Salas, J., et al, 2019, Wild Plants Potentially Used in Human Food in the Protected Area “Sierra Grande de Hornachos” of Extremadura (Spain). Sustainability 2019, 11, 456
  • Bodkin, F., 1991, Encyclopedia Botanica. Cornstalk publishing, p 1001
  • Boesi, A., 2014, Traditional knowledge of wild food plants in a few Tibetan communities. Journal of Ethnobiology and Ethnomedicine 10:75
  • Bonet, M. A. & Valles, J., 2002, Use of non-crop food vascular plants in Montseny biosphere reserve (Catalonia, Iberian Peninsula). International Journal of Food Sciences and Nutrition (2002) 53, 225–248
  • Bremness, L., 1994, Herbs. Collins Eyewitness Handbooks. Harper Collins. p 224
  • Bowser, M., 2017, Handout on Edible Plants of the Kenai Peninsula. USFWS Kenai National Wildlife Refuge p 19
  • Bussman, R. W., et al, 2016, A comparative ethnobotany of Khevsureti, Samtskhe-Javakheti, Tusheti, Svaneti, and Racha-Lechkhumi, Republic of Georgia (Sakartvelo), Caucasus. Journal of Ethnobiology and Ethnomedicine.
  • Bussman, R. W., et al, 2021, Unity in diversity—food plants and fungi of Sakartvelo (Republic of Georgia), Caucasus. Journal of Ethnobiology and Ethnomedicine (2021) 17:72 p 15
  • Cakir, E. A., 2017, Traditional knowledge of wild edible plants of Iğdır Province (East Anatolia, Turkey). Acta Soc Bot Pol. 2017;86(4):3568
  • Cerne, M., 1992, Wild Plants from Slovenia used as Vegetables. Acta Horticulturae 318.
  • Chen, W., et al, 2021, Wild plants used by the Lhoba people in Douyu Village, characterized by high mountains and valleys, in southeastern Tibet, China. Journal of Ethnobiology and Ethnomedicine (2021) 17:46
  • Chettri, N. & Sharma, E., Non-timber Forest Produce: Utilization, Distribution and Status in the Khangchendzonga Biosphere Reserve, Sikkim, India.
  • Ciocarlan, N. & Ghendov, V., 2015, Ethnobotanical and Ecological Studies of Wild Edible Plants from Bugeac Steppe, Republic of Moldova. Journal of EcoAgriTourism. Cailta terra Vol. 11(2):
  • Cordero, S. E., Abello, L. A., & Galvez, F. L., 2017, Plantas silvestres comestibles y medicinales de Chile y otras partes del mundo. CORMA p 252
  • Cundall, P., (ed.), 2004, Gardening Australia: flora: the gardener's bible. ABC Books. p 1451
  • Dangol, D. R. et al, 2017, Wild Edible Plants in Nepal. Proceedings of 2nd National Workshop on CUAOGR, 2017.
  • Demir, E., et al, 2020, Nutrient and bioactive substance contents of edible plants grown naturally in Salıpazarı (Samsun). Acta Sci. Pol. Hortorum Cultus, 19(1), 151–160
  • Demir, I. & Ayaz, N., 2022, Wild edible plants contributing to the traditional foods of Mardin (Turkey) Province. Indian Journal of Traditional Knowledge Vol 21(3), July 2022, pp 569-582
  • Demir, I. & Ayaz, N., 2022, Wild edible plants contributing to the traditional foods of Mardin (Turkey) Province. Indian Journal of Traditional Knowledge Vol 21(3), July 2022, pp 569-582 (As Urtica haussknechtii)
  • Denes, A., et al, 2012, Wild plants used for food by Hungarian ethnic groups living in the Carpathian Basin. Acta Societatis Botanicorum Poloniae 81 (4): 381-396
  • Dogan, A. & Tuzlaci, E., 2015, Wild Edible Plants of Pertek (Tunceli-Turkey). Marmara Pharmaceutical Journal 19: 126-135
  • Dogan, Y., 2012, Traditionally used wild edible greens in the Aegean Region of Turkey. Acta Societatis Botanicorum Poloniae 81(4): 329-342
  • Dogan, Y., et al, 2015, Of the importance of a leaf: the ethnobotany of sarma in Turkey and the Balkans. Journal of Ethnobiology and Ethnomedicine, 11:56
  • Dolina, K. & Luczaj, L., 2014, Wild food plants used on the Dubrovnik coast (south-eastern Croatia) Acta Soc Bot Pol 83(3):175–181
  • Duke, J.A., 1992, Handbook of Edible Weeds. CRC Press. p 204
  • Dutt, B., et al, 2011, Ethnobotanical Survey of Plants used by Gaddi Tribe of Bharmour Area in Himachal Pradesh. ENVIS Bulletin: Himalayan Ecology 19, 2011
  • Elias, T.S. & Dykeman P.A., 1990, Edible Wild Plants. A North American Field guide. Sterling, New York p 79
  • Ertug, F., 2000, An Ethnobotanical Study in Central Anatolia (Turkey). Economic Botany Vol. 54. No. 2. pp. 155-182
  • Ertug, F., 2004, Wild Edible Plants of the Bodrum Area. (Mugla, Turkey). Turk. J. Bot. 28 (2004): 161-174
  • Ertug, F, Yenen Bitkiler. Resimli Türkiye Florası -I- Flora of Turkey - Ethnobotany supplement
  • Esperanca, M. J., 1988. Surviving in the wild. A glance at the wild plants and their uses. Vol. 2. p 107
  • Facciola, S., 1998, Cornucopia 2: a Source Book of Edible Plants. Kampong Publications, p 243
  • Fox, F. W. & Young, M. E. N., 1982, Food from the Veld. Delta Books. p 359
  • Geraci, A., et al, 2018, The wild taxa utilized as vegetables in Sicily (Italy): a traditional component of the Mediterranean diet. Journal of Ethnobiology and Ethnomedicine (2018) 14:14
  • Ghanimi, R., et al, 2022, Ethnobotanical study on wild edible plants traditionally used by Messiwa people, Morocco. Journal of Ethnobiology and Ethnomedicine. 18:16
  • Ghimeray, A. K., Lamsal, K., et al, 2010, Wild edible angiospermic plants of the Illam Hills (Eastern Nepal) and their mode of use by local community. Korean J. Pl. Taxon. 40(1)
  • Ghimire, S. K., et al, 2008, Non-Timber Forest Products of Nepal Himalaya. WWF Nepal p 153
  • Ghosh, C. & Das A. P., 2011, Some useful and poisonous tea garden weeds from the Darjiling District of West Bengal, India. Pleione 5(1): 91 - 114
  • Girard, N. J., 2020, Sustainable Foraging of Wild Edible Plants in Norway. A Biocultural Approach. M. Sc. thesis Norwegian University. p 135
  • Guil-Guerrero, J. L., et al, 2001, Edible Wild Plants. in Recent Progress in Medicinal Plants Vol. 8 Sci. Tech publishing, Texas
  • Gunes, S. et al, 2018, Survey of wild food plants for human consumption in Karaisali (Adana-Turkey). Indian Journal of Traditional Knowledge. Vol. 17(2), April 2018, pp 290-298
  • GUPTA,
  • Hadjichambis, A. C., et al, 2007, Wild and semi-domesticated food plant consumption in seven circum-Mediterranean areas. International Journal of Food Sciences and Nutrition. 2007, 1-32.
  • Hammer, K. & Spahillari, M., 1999, Crops of European origin. in Report of a networking group on minor crops. IPGRI p 47
  • Hancer, C. K., et al, 2020, Traditional Knowledge of Wild Edible Plants of Biga (Çanakkale), Turkey. Acta Societatis Botanicorum Poloniae / 2020 / Volume 89 / Issue 1 / Article 8914
  • Hedrick, U.P., 1919, (Ed.), Sturtevant's edible plants of the world. p 665
  • Hibbert, M., 2002, The Aussie Plant Finder 2002, Florilegium. p 304
  • Hovsepyan, R., et al, 2016, Food as a marker for economy and part of identity: traditional vegetal food of Yezidis and Kurds in Armenia. Journal of Ethnic Foods. 3:32-41
  • http://nordicfood lab/org/blog/2102/9/wild-edible-plants-an-overview
  • http://www.botanic-gardens-ljubljana.com/en/plants
  • Irving, M., 2009, The Forager Handbook, A Guide to the Edible Plants of Britain. Ebury Press p 224
  • Jabeen, A., et al, 2009, Indigenous uses of economically important flora of Margallah Hills National Park, Islamabad, Pakistan. African Journal of Biotechnology Vol. 8 (5), pp. 763-784
  • Jardin, C., 1970, List of Foods Used In Africa, FAO Nutrition Information Document Series No 2.p 108
  • Joshi, N., et al, 2007, Traditional neglected vegetables of Nepal: Their sustainable utilization for meeting human needs. Tropentag 2007. Conference on International Agricultural Research for Development.
  • Joshi, N. & Siwakoti, M., 2012, Wild Vegetables Used by Local Community of Makawanpur District and Their Contribution to Food Security and Income Generation. Nepal Journal of Science and Technology Vol. 13, No. 1 (2012) 59-66
  • Kang, J. et al, 2016, Wild food plants and fungi used in the mycophilous Tibetan community of Zhagana (Tewo Country, Gansu, China) Journal of Ethnobiology and Ethnomedicine. 12:21
  • Kargioglu, M., et al, 2008, An Ethnobotanical Survey of Inner-West Anatolia, Turkey. Human Ecology 36:763-777
  • Kargioglu, M. et al, 2010, Traditional Uses of Wild Plants in the Middle Aegean Region. Human Ecology 38:429-450
  • Kaya, O. M., et al, 2020, An ethnobotanical research in Sanhurfa central district and attached Villages (Turkey). Indian Journal of Traditional Knowledge. Vol. 19(1) pp 7-23
  • Kayabasi, N. P., et al, 2018, Wild edible plants and their traditional use in the human nutrition in Manyas (Turkey). Indian Journal of Traditional Knowledge. Vol. 17(2), April 2018, pp 299-306
  • Kays, S. J., and Dias, J. C. S., 1995, Common Names of Commercially Cultivated Vegetables of the World in 15 languages. Economic Botany, Vol. 49, No. 2, pp. 115-152
  • Kermath, B. M., et al, 2014, Food Plants in the Americas: A survey of the domesticated, cultivated and wild plants used for Human food in North, Central and South America and the Caribbean. On line draft. p 889
  • Khakurel, D., et al, 2021, Foods from the wild: Local knowledge, use pattern and distribution in Western Nepal. PLOS ONE.
  • Khan, A. H., et al, 2023, Traditional foraging for ecological transition? Wild food ethnobotany among three ethnic groups in the highlands of the eastern Hindukush, North Pakistan. Journal of Ethnobiology and Ethnomedicine 19:9
  • Khan, M. T., et al, 2016, Ethnobotanical Study of Wild Flora in the Remote Areas of Northern Pakistan. Wulfenia Journal Vol. 23, No 10 p 161
  • Khojimatov, O. K. et al, 2015, Some wild growing plants in traditional foods of Uzbekistan. Journal of Ethnic Foods. 2 (2015) 25-28
  • Kiple, K.F. & Ornelas, K.C., (eds), 2000, The Cambridge World History of Food. CUP p 1822
  • Kizilarslan, C. & Ozhatay, N., 2012, An ethnobotanical study of the useful and edible plants of İzmit. Marmara Pharmaceutical Journal 16: 134-140, 2012.
  • Koca, I., et al, 2015, Some Wild Edible Plants and Their Dietary Fiber Contents. Pakistan Journal of Nutrition. 14(4): 188-194
  • Kolosova, V., et al, 2020, Foraging in Boreal Forest: Wild Food Plants of the Republic of Karelia, NW Russia. Foods 2020, 9, 1015; p 20
  • Kumar, A., et al, 2012, Ethnobotanical Edible Plant Biodiversity of Lepcha Tribes. Indian Forester, 138 (9):798-803
  • Kybal, J., 1980, Herbs and Spices, A Hamlyn Colour Guide, Hamlyn Sydney p 200
  • Lamp, C & Collet F., 1989, Field Guide to Weeds in Australia. Inkata Press. p 317
  • Letsela, T., et al, 2003, Plant Resources Used for Subsistence in Tsehlanyane and Boking in Lesotho. Economic Botany 57(4): 619-639
  • Low, T., 1991, Wild Herbs of Australia and New Zealand. Angus & Robertson. p 93
  • Luczaj, L., 2012, Ethnobotanical review of wild edible plants of Slovakia. Acta Societatis Botanicorum Poloniae 81(4):245-255
  • Luczaj, L. et al, 2012, Wild food plant use in 21st century Europe: the disappearance of old traditions and the search for new cuisines involving wild edibles. Acta Soc Bot Pol 81(4):359–370
  • Luczaj, L. et al, 2013, Wild edible plants of Belarus: from Rostakinski's questionnaire of 1883 to the present. Journal of Ethnobiology and Ethnomedicine 9:21
  • Luczaj, L. et al, 2013, Wild food plants used in the villages of the Lake Vrana Nature Park (northern Dalmatia, Croatia). Acta Societatis Botanicorum Poloniae, 82(4): 275-281
  • Luczaj, L. et al, 2014, Wild Food Plants of Dalmatia (Croatia). in A. Pieroni, C. L. Quave (eds.), Ethnobotany and Biocultural Diversities in the Balkans, p 143
  • Łukasz Łuczaj and Wojciech M Szymański, 2007, Wild vascular plants gathered for consumption in the Polish countryside: a review. J Ethnobiol Ethnomedicine. 3: 17
  • Luczaj, L., et al, 2015, Wild food plants and fungi used by Ukrainians in the western part of the Maramureş region in Romania. Acta Soc Bot Pol 84(3):339–346
  • Luczaj, L. et al, 2017, Comfrey and Buttercup Eaters: Wild Vegetables of the Imereti Region in Western Georgia, Caucasus. Economic Botany, 71(2), 2017, pp. 188–193
  • Mabey, R., 1973, Food for Free. A Guide to the edible wild plants of Britain, Collins. p 100
  • MacKinnon, A., et al, 2009, Edible & Medicinal Plants of Canada. Lone Pine. p 312
  • Magwede, K., van Wyk, B.-E., & van Wyk, A. E., 2019, An inventory of Vhavenḓa useful plants. South African Journal of Botany 122 (2019) 57–89
  • Mahklouf, M. H., 2019, Ethnobotanical Study of Edible Wild Plants in Libya. European Journal of Ecology. 5(2): 30-40
  • Manandhar, N.P., 2002, Plants and People of Nepal. Timber Press. Portland, Oregon. p 469
  • Martin, F.W. & Ruberte, R.M., 1979, Edible Leaves of the Tropics. Antillian College Press, Mayaguez, Puerto Rico. p 223
  • Masoodi, H. U. R. & Sundriyal, R. C., 2020, Richness of non-timber forest products in Himalayan communities—diversity, distribution, use pattern and conservation status. Journal of Ethnobiology and Ethnomedicine 16:56
  • Mattalia, G., et al, 2012, Traditional uses of wild food and medicinal plants among Brigasc, Kye ́, and Provenc ̧al communities on the Western Italian Alps. Genet Resour Crop Evol. Springer
  • Michael, P., 2007, Edible Wild Plants and Herbs. Grub Street. London. p 227
  • Mirdeilami, S. Z., et al, 2011, Ethnopharmacological survey of medicinal plants in Maraveh Tappeh region, north of Iran. Iranian Journal of Plant Physiology 2(1): 327‐338
  • Miskoska-Milevska, E. et al, 2020, Traditional uses of wild edible plants in the Republic of North Macedonia. PHYTOLOGIA BALCANICA 26(1): 155–162, Sofia, 2020 p 159
  • Miya, M. S. & Gautum, D., 2021, Checklist of floral species at the Institute of Forestry, Pokhara Campus, Pokhara, Nepal. Journal of Institute of Forestry, Nepal 18 (2021) 133-177
  • Moerman, D. F., 2010, Native American Ethnobotany. Timber Press. p 579
  • Mukemre, M., et al, 2016, Survey of wild food plants for human consumption in villages of Catak, (Van-Turkey), Indian Journal of Traditional Knowledge. Vol. 15(2) pp. 183-191
  • Nanagulyan, S., et al, 2020, Wild plants and fungi sold in the markets of Yerevan (Armenia). Journal of Ethnobiology and Ethnomedicine 16:26
  • Nassif, F., & Tanji, A., 2013, Gathered food plants in Morocco: The long forgotten species in Ethnobotanical Research. Life Science Leaflets 3:17-54
  • Nedelcheva A., 2013, An ethnobotanical study of wild edible plants in Bulgaria. EurAsian Journal of BioSciences 7, 77-94
  • Özdemir, E. and Kültür, S., 2017, Wild Edible Plants of Savaştepe District (Balıkesir, Turkey), Marmara Pharm J 21/3: 578-589
  • Paoletti, M.G., Dreon, A.L., and Lorenzoni, G.G., 1995, Pistic, Traditional Food from Western Friuli, NE Italy. Economic Botany 49(1) pp 26-30
  • Pardo-de-Santayana, M., et al, 2005, The gathering and consumption of wild edible plants in the Campoo (Cantabria, Spain). International Journal of Food Sciences and Nutrition. 56(7): 529-542
  • Pardo-de-Santayana, M., et al, 2007, Traditional knowledge of wild edible plants used in the northwest of the Iberian Peninsula (Spain and Portugal): a comparative study. Journal of Ethnobiology and Ethnomedicine 2007, 3:27
  • Pasta, S., et al, 2020, An Updated Checklist of the Sicilian Native Edible Plants: Preserving the Traditional Ecological Knowledge of Century-Old Agro-Pastoral Landscapes. Frontiers in Plant Science. Volume 11|Article 388
  • Pieroni, A., 1999, Gathered wild food plants in the Upper Valley of the Serchio River (Garfagnana), Central Italy. Economic Botany 53(3) pp 327-341
  • Pieroni, A.,& Giusti, M. E., 2009, Alpine Ethnobotany in Italy: Journal of Ethnobiology and Ethnomedicine. 5:32
  • Pieroni, A., 2008, Local plant resources in the ethnobotany of Theth, a village in the Northern Albanian Alps. Genet Resour Crop Evol (2008) 55:1197–1214
  • Pieroni, A., et al, 2012, "We are Italians!": The Hybrid Ethnobotany of a Venetian Diaspora in Eastern Romania. Human Ecology 40:435-451
  • Pieroni, A. et al, 2014, Resilience at the border: traditional botanical knowledge among Macedonians and Albanian living in Gollobordo, Eastern Albania. Journal of Ethnobiology and Ethnomedicine. 10:31
  • Pieroni, A. & Soukand, R., 2017, Are Borders more Important than Geographical Distance? The Wild Food Ethnobotany of the Boykos and its Overlap with that of the Bukovinian Hutsuls in Western Ukraine. Journal of Ethnobiology 37(2): 326–345
  • Pieroni, A. & Soukand, R., 2017, The disappearing wild food and medicinal plant knowledge in a few mountain villages of North-Eastern Albania. Journal of Applied Botany and Food Quality 90, 58 - 67
  • Pieroni, A. & Soukand, R., 2018, Forest as Stronghold of Local Ecological Practice: Currently Used Wild Food Plants in Polesia, Northern Ukraine. Economic Botany, XX(X) pp. 1-21
  • Pieroni, A., et al, 2020, Wild food plants traditionally gathered in central Armenia: archaic ingredients or future sustainable foods? Environment, Development and Sustainability. Springer p 12
  • Plants for a Future database, The Field, Penpol, Lostwithiel, Cornwall, PL22 0NG, UK. http://www.scs.leeds.ac.uk/pfaf/
  • Polat, R., et al, 2015, Survey of wild food plants for human consumption in Elazig (Turkey). Indian Journal of Traditional Knowledge. Vol. 1(1): 69-75
  • Polat, R., et al, 2017, Survey of wild food plants for human consumption in Bingol, (Turkey). Indian Journal of Traditional Knowledge. Vol. 16(3) July 2017, pp. 378-384
  • Polunin, O., & Stainton, A., 2006, Flowers of the Himalaya, Oxford India Paperbacks. p 366
  • Prakofjewa, J., et al, 2023, Boundaries Are Blurred: Wild Food Plant Knowledge Circulation across the Polish-Lithuanian-Belarusian Borderland. Biology 2023, 12, 571.
  • Pruse, B., et al, 2021, Active Wild Food Practices among Culturally Diverse Groups in the 21st Century across Latgale, Latvia. Biology 2021, 10, 551.
  • Radha, B., et al, 2013, Wild Edible Plant Resources of the Lohba Range of Kedarnath Forest Division (KFD), Garhwal Himalaya, India. Int. Res J. Biological Sci. Vol. 2 (11), 65-73
  • Rana, D., et al, 2019, Ethnobotanical knowledge among the semi-pastoral Gujjar tribe in the high altitude (Adhwari’s) of Churah subdivision, district Chamba, Western Himalaya. Journal of Ethnobiology and Ethnomedicine (2019) 15:10
  • Rana, D., et al, 2020, Use of medicinal plants for treating different ailments by the indigenous people of Churah subdivision of district Chamba, Himachal Pradesh, India. Environment, Development and Sustainability. Springer.
  • Rana, J.C. et al, 2011, Genetic resources of wild edible plants and their uses among tribal communities of cold arid regions of India. Genetic Resources and Crop Evolution. 59:135-149
  • Rana, P. K., et al, 2014, Uses of Local Plant Biodiversity among the Tribal Communities of Pangi Valley of District Chamba in Cold Desert Himalaya, India. The Scientific World Journal. Volume 2014, Article ID 753289, 15 pages
  • Redzic, S. J., 2006, Wild Edible Plants and their Traditional Use in the Human Nutrition in Bosnia-Herzegovina. Ecology of Food and Nutrition, 45:189-232
  • Redzic, S., 2010, Use of Wild and Semi-Wild Edible Plants in Nutrition and Survival of People in 1430 Days of Siege of Sarajevo during the War in Bosnia and Herzegovina (1992–1995). Coll. Antropol 34 (2010) 2:551-570
  • Rexhepi, B., et al, 2018, Ethnobotanical Study of Wild Edible Plants in Pelagonia Region (Southwestern Macedonia) International Journal of Advances in Science Engineering and Technology, Vol. 6 (1) p 57
  • Rigat, M et al, 2009, Ethnobotany of Food Plants in the High River Ter Valley (Pyrenees, catalonia, Iberian Peninsula): Non-Crop Food Vascular Plants and Crop Food Plants with medicinal Properties. Ecology of Food and Nutrition, 48:303-327
  • Rivera, C., et al, 2005, The Ethnobotanical Study of Local Mediterranean Food Plants as Medicinal Resources in Southern Spain. Journal of Physiologyand Pharmacology. 56, Suppl. 1, 97-114
  • Ruiters-Welcome, A. K., 2019, Food plants of southern Africa. Ph.D. thesis. Univ. of Johannesburg p 106
  • Salvi, J. et al, 2016, A review: Underutilized wild edible plants as a potential source of alternative nutrition. International Journal of Botany Studies. Volume 1; Issue 4; May 2016; Page No. 32-36
  • Sansanelli, S., et al, 2014, Wild food plants traditionall consumed in the area of Bologna (Emilia Romagna region, Italy). Journal of Ethnobiology and Ethnomedicine 10:69
  • Sansanelli, S., et al, 2017, Ethnobotanical survey of wild food plants traditionally collected and consumed in the Middle Agri Valley (Basilicata region, southern Italy). Journal of Ethnobiology and Ethnomedicine (2017) 13:50
  • Saunders, C.F., 1948, Edible and Useful Wild Plants. Dover. New York. p 127
  • Savita, et al, 2006, Studies on wild edible plants of ethnic people in east Sikkim. Asian J. of Bio Sci. (2006) Vol. 1 No. 2 : 117-125
  • Schneider, E., 2001, Vegetables from Amaranth to Zucchini: The essential reference. HarperCollins. p 417
  • Schunko, C., et al, 2010, Organic farmers use of wild food plants and fungi in a hilly area in Styria (Austria). Journal of Ethnobiology and Ethnomedicine 6:17
  • Schunko, C., et al, 2012, Intracultural variation of knowledge about wild plant uses in the Biosphere Reserve Grosses Walsertal (Austria). Journal or Ethnobiology and Ethnomedicine, 8:23
  • Senkardes, I & Tuzlaci, E., 2016, Wild Edible Plants of Southern Part of Nevsehir inTurkey. Marmara Pharmaceutical Journal 20:34-43
  • Sharma, L. et al, 2018, Diversity, distribution pattern, endemism and indigenous uses of wild edible plants in Cold Desert Biosphere Reserve of Indian Trans Himalaya. Indian Journal of Traditional Knowledge. Vol 17(1) January 2018 pp 122-131
  • Sharma, P., et al, 2013, Wild edibles of Murari Devi and surrounding areas in Mandi district of Himachal Pradesh, India. International Journal of Biodiversity and Conservation. Vol. 5(9), pp. 580-592, September 2013
  • Kumar, P. D., et al, 2015, Ethnobotanical Knowledge and Usage of Wild Plants in Theog Forest Division, Himachal Pradesh, North Western Himalaya. The Journal of Ethnobiology and Traditional Medicine. Photon 124(2015) 922-935
  • Shava, S., 2000, The Use of Indigenous Plants as Food by a Rural Community in the Eastern Cape: an Educational Exploration. Masters Thesis Rhodes University. p 67
  • Sher, H. et al, 2011, Ethnobotanical and Economic Observations of Some Plant Resources from the Northern Parts of Pakistan. Ethnobotany research & Applications 9:027-041
  • Shikov, A. N. et al, 2017, Traditional and Current Food Use of Wild Plants Listed in the Russian Pharmacopoeia. Frontiers in Pharmacology. Vol. 8 Article 841
  • Siminiuc, R. & Turcanu, D., 2021, Study of Edible Spontaneous Herbs in the Republic of Moldova for Ensuring a Sustainable Food System. Food and Nutrition Sciences, 2021, 12, 703-718 p 709
  • Simkova, K. et al, 2014, Ethnobotanical review of wild edible plants used in the Czech Republic. Journal of Applied Botany and Food Quality 88, 49-67
  • Singh, A. G., Panthi, M. P., & Tewari, D. D., 2012, Wild Plants Used as Vegetable in Rupandehi District of Nepal and their Ethnomedicinal Importance. J. Nat. Hist. Mus. Vol. 26, 2012, 111-125
  • Singh, H.B., Arora R.K.,1978, Wild edible Plants of India. Indian Council of Agricultural Research, New Delhi. p 38
  • Skinner, G. & Brown, 1981, C., Simply Living. A gatherer's guide to New Zealand's fields, forests and shores. Reed. p 15
  • Soukand, R., et al, 2017, Multi-functionality of the few: current and past uses of wild plants for food and healing in Liubań region, Belarus. Journal of Ethnobiology and Ethnomedicine (2017) 13:10
  • Soukand, R., et al, 2021, The trauma of no-choice: Wild food ethnobotany in Yaghnobi and Tajik villages, Varzob Valley, Tajikistan. Genetic Resources and Crop Evolution · December 2021
  • Sp. pl. 2:984. 1753
  • Stryamets, N., et al, 2015, From economic survival to recreation: contemporary uses of wild food and medicine in rural Sweden, Ukraine and NW Russia. Journal of Ethnobiology and Ethnomedicine 11:53
  • Sundriyal, M., et al, 1998, Wild edibles and other useful plants from the Sikkim Himalaya, India. Oecologia Montana 7:43-54
  • Sundriyal, M., et al, 2004, Dietary Use of Wild Plant Resources in the Sikkim Himalaya, India. Economic Botany 58(4) pp 626-638
  • Svanberg, I., 2012, The use of wild plants as food in pre-industrial Sweden. Acta Societatis Botanicorum Poloniae 81(4): 317-327
  • Tardio, J., et al, Ethnobotanical review of wild edible plants in Spain. Botanical J. Linnean Soc. 152 (2006), 27-71
  • Tbatou, M, et al, 2016, Wild Edible Plants traditionally used in the countryside of El Jadida, Coastal Area in the Center of Morocco. Life Sciences Leaflets 75:28-48
  • Thakur, D., et al, 2017, Why they eat, what they eat: patterns of wild edible plants consumption in a tribal area of Western Himalaya. Journal of Ethnobiology and Ethnomedicine (2017) 13:70
  • Thapa, L. B., et al, 2014, Wild Edible Plants used by endangered and Indigenous Raji Tribe in Western Nepal. International Journal of Applied Sciences and Biotechnology. Vol 2(3):243-252
  • Tsering, J., et al, 2017, Ethnobotanical appraisal on wild edible plants used by the Monpa community of Arunchal Pradesh. Indian Journal of Traditional Knowledge. Vol 16(4), October 2017, pp 626-637
  • Tsherig, K., 2012, Edible Wild Plants of Bhutan and their contribution to Food and Nutrition Security. Ministry of Ag. and Forests, Bhutan. www.fao.org
  • Turner, N., 1995, Food Plants of Coastal First Peoples. Royal BC Museum Handbook p 131
  • Turner, N., et al, 2011, "Up on the Mountain": Ethnobotanical Important of Montane Sites in Pacific Coastal North America. Journal of Ethnobiology 31(1): 4-43
  • Ulian, T., et al, 2020, Unlocking plant resources to support food security and promote sustainable agriculture. Plants, People, Planet. 2020;2:421–445.
  • Uprety, Y., et al, 2011, Plant biodiversity and ethnobotany inside the projected impact area of the Upper Seti Hydropower Project, Western Nepal. Environ. Dev. Sustain. (2011) 13:463-492
  • Uprety, Y., et al, 2012, Diversity of use and local knowledge of wild edible plant resources in Nepal. Journal of Ethnobotany and Ethnomedicine 8:16
  • van Wyk, B., 2005, Food Plants of the World. An illustrated guide. Timber press. p 372
  • Vorstenbosch, T., et al, 2017, Famine food of vegetal origin consumed in the Netherlands during World War II. Journal of Ethnobiology and Ethnomedicine (2017) 13:63
  • Welcome, A. K. & Van Wyk, B.-E., 2019, An inventory and analysis of the food plants of southern Africa. South African Journal of Botany 122 (2019) 136–179
  • Wild Edible Plants in Lebanon. Promoting dietary diversity in poor communities in Lebanon. http://www.wildedibleplants.org/
  • World Checklist of Useful Plant Species 2020. Royal Botanic Gardens, Kew
  • www.wildediblefood.com
  • Yesil, Y., et al, 2019, Wild edible plants in Yeşilli (Mardin-Turkey), a multicultural area. Journal of Ethnobiology and Ethnomedicine (2019) 15:52
  • Yesil, Y., et al, 2019, Traditional knowledge of wild edible plants in Hasankeyf (Batman Province, Turkey). Acta Societatis Botanicorum Poloniae. 88(3):3633

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